The taxonomy of the leafy liverwort genus Leptoscyphus (Lophocoleaceae) revisited

Abstract

A revised classification of the fiverwort genus Leptoscyphus (Lophocoleaccac) is presented on the basis of previous phylogenetic analyses as well as morphological examination of recent plain material and type specimens. The three subgenera that have traditionally been recognized based on morphological.,rounds, namely Leptoscyphus subg. Anomylia, L subg. Physoscyphus, and L. subg. Leoptoscyphus, are well retrieved by the phylogeny. Two new subgencra, L. subg. Spinoscyphus and L. subg. Australoleptoscyphus, are created I-or L. cleefii and L. spectabilis and the Southern South Hemisphere species, respectively. Three new species, namely L. gradsteinii, L. lainhinonii, and L. solimixii, arc described here based on molecular and rnorphological evidence. The new combination L. porphyrius subsp. azoricus is made. Lectotypcs are designated for Chiloscyphus australis, C. huidobroamus, Jungermannia amphibolia, J pallidovirens J. physocalyx Lophocolea behnoi-ana, L. diversi/blia, L. excipulata, and Mylia azorica